Last updated on 06 December, 2024

Record statistics by species and TCR chain

General statistics. Note that general statistics are computed using the ‘slim’ database version. Slim version, for example, lists the same TCR sequence found in several donors/studies only once and selects representative V/J for a given CDR3aa clonotype and the best score across all redundant records.

Species Chain Records Paired records Unique epitopes Studies
HomoSapiens TRA 33170 33170 1089 210
HomoSapiens TRB 48064 28156 1279 264
MacacaMulatta TRB 1279 0 3 2
MusMusculus TRA 1694 1694 57 36
MusMusculus TRB 2825 1641 79 40

Record statistics by year

Number of unique TCR sequences, epitopes, MHC alleles and studies by publication year, cumulative plots.

Summary by antigen and antigen origin

Representative data for Homo Sapiens

Parent species Records Unique epitopes Studies
CMV 33192 31 49
InfluenzaA 11802 30 29
EBV 10963 33 37
SARS-CoV-2 8986 671 18
HomoSapiens 7443 330 108
HIV-1 3248 64 34
HCV 1889 22 11
Mtb 816 2 1
YFV 677 1 5
DENV 399 3 1
Influenza B 377 3 1
PlasmodiumFalciparum 282 6 1
TriticumAestivum 233 19 12
HTLV-1 217 8 9
SARS-CoV 143 1 1
Wheat 86 2 3
RotavirusA 84 3 1
CoxsackievirusB 65 2 1
synthetic 62 29 4
M.tuberculosis 61 11 3
HCoV-HKU1 48 1 1
HSV-2 47 1 1
E.Coli 41 3 4
HPV 28 1 1
AdV 15 2 1
MCPyV 12 2 1
StreptomycesKanamyceticus 6 1 1
HPV-16 4 2 1
HHV 2 1 1
PseudomonasAeruginosa 2 1 1
PseudomonasFluorescens 2 1 1
SaccharomycesCerevisiae 2 1 1

COVID-19 data

Summary of antigens and T-cell receptors related to COVID-19 pandemic. Number of records for SARS-CoV-2 epitopes grouped by viral protein and HLA plotted using alluvium plot. Epitopes with less than 30 records in total were not counted.

Summary of SARS-CoV-2 epitopes and corresponding TCR alpha and beta chain specificity records (cases with 10+ records)

Gene HLA Epitope Studies TRA TRB
Spike HLA-A.02 YLQPRTFLL 9 678 903
Nucleocapsid HLA-B.07 SPRWYFYYL 4 386 544
ORF1ab HLA-A.01 TTDPSFLGRY 3 383 438
Spike HLA-DPA.01/B.04 TFEYVSQPFLMDLE 2 100 444
Nucleocapsid HLA-A.03 KTFPPTEPK 2 128 167
Spike HLA-A.24 QYIKWPWYI 5 137 150
Spike HLA-A.01 LTDEMIAQY 2 125 129
Spike HLA-B.15 NQKLIANQF 1 75 73
Nucleocapsid HLA-A.02 RTATKQYNV 1 50 93
Spike HLA-A.02 RLQSLQTYV 5 53 88
Nucleocapsid HLA-B.40 MEVTPSGTWL 1 46 62
Spike HLA-A.24 NYNYLYRLF 3 56 52
ORF3 HLA-A.01 FTSDYYQLY 2 41 40
ORF3 HLA-A.02 LLYDANYFL 3 28 41
ORF1ab HLA-A.02 ALWEIQQVV 2 27 36
ORF3 HLA-A.02 ALSKGVHFV 3 20 40
ORF3 HLA-A.24 VYFLQSINF 2 29 29
Matrix HLA-B.15 RVAGDSGFAAY 1 24 24
ORF1ab HLA-A.02 KLWAQCVQL 1 21 25
Nucleocapsid HLA-A.02 RTATKAYNV 1 19 23
ORF1ab HLA-A.01 PTDNYITTY 2 16 26
Matrix HLA-B.07 WPVTLACFV 1 14 21
ORF1ab HLA-A.02 KSVNITFEL 1 16 19
Spike HLA-B.07 SIIAYTMSL 1 15 17
Spike HLA-DRA.01/B1.04 HWFVTQRNFYEPQII 1 16 16
ORF1ab HLA-A.02 VMVELVAEL 1 13 17
Matrix HLA-A.02 WLLWPVTLA 1 9 16
NSP3 HLA-A.01 PTDNYITTY 1 5 19
ORF14 HLA-A.02 LLLEWLAMA 1 10 13
Nucleocapsid HLA-A.02 LLLDRLNQL 2 4 18
ORF1ab HLA-A.02 TLMNVLTLV 1 9 13
Spike HLA-DRA.01/B1.04 NCTFEYVSQPFLMDL 1 6 16
ORF1ab HLA-A.24 TSQWLTNIF 1 10 11
ORF9b HLA-A.02 KVYPIILRL 1 9 12
Spike HLA-B.07 WPWYIWLGF 1 7 14
Spike HLA-DRA.01/B1.11 VGGNYNYLYRLFRKS 1 10 10
ORF1ab HLA-A.02 CLAVHECFV 1 9 10
ORF1ab HLA-A.02 FLAHIQWMV 1 8 11
ORF1ab HLA-A.02 FLLNKEMYL 1 9 10
ORF1ab HLA-A.02 RQLLFVVEV 1 9 10
NSP12 HLA-A.24 VYIGDPAQL 2 9 9
Nucleocapsid HLA-B.44 QELIRQGTDY 1 9 9
ORF1ab HLA-A.02 AIFYLITPV 1 7 11
ORF1ab HLA-A.02 FTVLCLTPV 1 7 11
Spike HLA-A.02 VFLVLLPLV 1 8 10
ORF1ab HLA-A.24 NYMPYFFTL 1 8 9
ORF3 HLA-A.02 YLYALVYFL 1 7 10
Spike HLA-A.02 RLITGRLQSL 1 9 8
ORF1ab HLA-A.01 NSSTCMMCY 1 6 10
ORF1ab HLA-A.02 KLQFTSLEI 1 8 8
ORF1ab HLA-A.02 YVDNSSLTI 1 7 9
ORF1ab HLA-A.02 KLKDCVMYA 1 7 8
ORF1ab HLA-A.02 YVWKSYVHV 1 6 9
ORF1ab HLA-B.07 MPASWVMRI 1 5 10
ORF1ab HLA-B.07 MPYFFTLLL 1 7 8
ORF7a HLA-B.07 SPIFRIVAA 1 4 11
Nucleocapsid HLA-A.01 NTNSSPDDQIGYY 1 7 7
Nucleocapsid HLA-B.07 RIRGGDGKM 1 6 8
ORF10 HLA-A.02 NVFAFPFTI 1 7 7
ORF1ab HLA-A.02 FLALCADSI 1 7 7
ORF1ab HLA-A.02 FLPRVFSAV 1 6 8
ORF1ab HLA-A.02 SVLYYQNNV 1 6 8
ORF1ab HLA-A.02 YMPYFFTLL 1 7 7
ORF1ab HLA-A.02 YTMADLVYA 1 5 9
RNP HLA-A.01 DTDFVNEFY 2 6 8
Spike HLA-A.02 IMLCCMTSC 1 6 8
Spike HLA-B.44 AEVQIDRLI 1 7 7
Spike HLA-DRA.01/B1.03 RISNCVADYSVLYNS 1 7 7
ORF1ab HLA-A.02 FLPGVYSVI 1 5 8
ORF1ab HLA-A.02 LLDDFVEII 1 6 7
Spike HLA-A.02 KQIYKTPPI 2 6 7
Spike HLA-DRA.01/B1.15 NLLLQYGSFCTQLNRAL 1 0 13
Matrix HLA-A.24 LWLLWPVTL 1 6 6
Nucleocapsid HLA-A.02 LLNKHIDAY 1 4 8
ORF1ab HLA-A.02 KLNEEIAII 1 5 7
ORF1ab HLA-A.02 KLSYGIATV 1 4 8
ORF1ab HLA-A.02 LMIERFVSL 1 5 7
ORF1ab HLA-A.02 RIMTWLDMV 1 4 8
ORF1ab HLA-A.02 SMMILSDDA 1 6 6
ORF1ab HLA-A.02 YLFDESGEF 1 5 7
ORF1ab HLA-B.07 SPYNSQNAV 1 4 8
Spike HLA-DRA.01/B1.03 NFSQILPDPSKPSKR 1 6 6
Matrix HLA-A.02 LACFVLAAV 1 4 7
ORF1ab HLA-A.01 YADVFHLYL 1 5 6
ORF1ab HLA-A.02 FLNRFTTTL 1 5 6
ORF1ab HLA-A.02 TLMNVITLV 1 5 6
ORF1ab HLA-A.02 TTIQTIVEV 1 5 6
ORF1ab HLA-A.02 YADVFHLYL 1 5 6
ORF1ab HLA-A.02 YLGGMSYYC 1 5 6
ORF1ab HLA-B.07 VPYFNMVYM 1 4 7
Spike HLA-A.02 FIAGLIAIV 2 4 7
ORF1ab HLA-A.01 FLTENLLLY 1 4 6
ORF1ab HLA-A.01 TSAMQTMLF 1 4 6
ORF1ab HLA-A.02 LMCQPILLL 1 4 6
ORF1ab HLA-A.02 MLDMYSVML 1 5 5
ORF1ab HLA-A.02 SMWALVISV 1 5 5
ORF1ab HLA-A.02 TLKNTVCTV 1 5 5
ORF1ab HLA-A.24 VMHANYIFW 1 5 5
ORF7a HLA-A.02 KLFIRQEEV 1 5 5
ORF7b HLA-A.02 LLFLVLIML 1 5 5
Spike HLA-A.02 FLHVTYVPA 1 5 5
Spike HLA-DQA1.01/B1.06 NNSYECDIPIGAGIC 1 5 5

Self-antigen data

Summary of T-cell receptors recognizing self-antigens, including antigens linked to utoimmune diseases and potential neoantigen targets for cancer immunotherapy. Number of records for self-antigens grouped by (mutated) human gene and corresponding HLAs are plotted using alluvium plot. Only self-antigens with at least 10 records are shown.

Summary of neoantigens and corresponding TCR alpha and beta chain specificity records (cases with 5+ records)

Gene HLA Epitope Studies TRA TRB
MLANA HLA-A.02 ELAGIGILTV 16 364 1434
PGT HLA-A.02 LLAGIGTVPI 1 434 882
BST2 HLA-A.02 LLLGIGILV 1 380 233
MART1 HLA-A.02 ALAGIGILTV 1 95 150
APOB HLA-DRA.01/B1.07 SLFFSAQPFEITAST 1 103 104
IGF2BP2 HLA-A.02 NLSALGIFST 1 0 112
MLANA HLA-A.02 ALGIGILTV 2 40 52
SF3B1 HLA-A.02 RLPGVLPRA 1 42 48
TKT HLA-A.02 AMFWSVPTV 1 0 83
G6PC2 HLA-A.01 LTSLTILQLY 1 34 36
ZNT8 HLA-B.08 YAKWKLCSA 1 28 41
PLA2G6 HLA-A.02 FLASKIGRLV 1 34 34
ZNT8 HLA-B.08 YAYAKWKL 1 31 32
SEC24A HLA-A.02 FLYNLLTRV 1 0 61
ZNT8 HLA-A.02 LLSLFSLWL 1 23 38
MLANA HLA-A.02 EAAGIGILTV 7 15 45
PMEL HLA-A.02 IMDQVPFSV 3 13 45
ZNT8 HLA-A.01 VTDAAHLLI 1 23 33
SLC30A8 HLA-A.02 VAANIVLTV 2 24 29
ZNT8 HLA-A.02 VVTGVLVYL 1 21 27
IAPP HLA-A.01 STNVGSNTY 1 16 27
INS HLA-DQA1.03/B1.02 GQVELGGGNAVEVCK 2 21 21
GFAP HLA-B.08 HLKRNIVV 1 16 23
GAD65 HLA-A.02 VMNILLQYV 1 14 24
G6PC2 HLA-B.08 NLIFKWIL 1 18 19
INS HLA-B.08 MALWMRLL 1 17 20
GAD2 HLA-B.08 MMIARFKM 1 15 21
PORCN HLA-A.02 LLHGFSFYL 1 0 36
IGRP HLA-A.02 FLWSVFWLI 1 12 22
MLANA HLA-A.02 ELAGIGLTV 1 20 13
PREINS HLA-A.02 ALWMRLLPL 1 15 18
PMEL HLA-A.02 KTWGQYWQV 1 16 16
PMEL HLA-A.02 YLEPGPVTV 2 1 31
ZNT8 HLA-A.02 IQATVMIIV 1 12 20
WT1 HLA-A.24 CYTWNQMNL 1 15 15
NY-ESO-1 HLA-DQA.01/B1.07 LLEFYLAMPFATP 1 13 16
ZNT8 HLA-A.02 KMYAFTLES 1 9 20
IGRP HLA-A.02 VLFGLGFAI 1 10 18
WT1 HLA-A.02 RMFPNAPYL 5 14 14
PREINS HLA-A.02 RLLPLLALL 1 12 15
G6PC2 HLA-B.08 YLKTNLFL 1 9 15
GAD2 HLA-A.01 VSATAGTTVY 1 14 10
IAPP HLA-B.08 ILKLQVFL 1 9 15
INS HLA-A.01 GSHLVEALY 1 10 14
INS HLA-A.02 RLLPLLALLAL 1 7 17
NY-ESO-1 HLA-A.02 SLLMWITQV 4 11 10
ZDBF2 HLA-A.02 YILKYSVFL 1 0 21
IAPP HLA-A.02 KLQVFLIVL 1 6 14
SRPX HLA-A.02 TLWCSPIKV 2 10 10
AKAP13 HLA-A.02 KLMNIQQKL 1 0 19
NY-ESO-1 HLA-DRA.01/B1.04 PGVLLKEFTVSGNIL 1 9 10
PTPRN HLA-A.02 LLPPLLEHL 1 8 9
ZNT8 HLA-A.02 ILSAHVATA 1 8 9
IGRP HLA-A.02 LNIDLLWSV 1 8 8
IGRP HLA-A.02 NLFLFLFAV 1 5 11
MAGEA1 HLA-A.02 KVLEYVIKV 1 8 8
ZNT8 HLA-A.02 LAVDGVLSV 1 9 7
ADH HLA-A.02 RQFGPDWIVA 1 6 9
FNDC3B HLA-A.02 VVMSWAPPV 1 7 8
GFAP HLA-B.08 LRLRLDQL 1 6 9
INS HLA-B.08 LWMRLLPL 1 7 8
MLANA HLA-A.02 AAGIGILTV 4 5 10
ZNT8 HLA-A.02 LLSILCIWV 1 5 10
5T4 HLA-A.02 RLARLALVL 1 7 7
BCL2L1 HLA-A.02 YLNDHLEPWI 1 7 7
INS HLA-B.08 WMRLLPLL 1 6 8
SLC3A2 HLA-DRA.01/B1.07 DPPALASTNAEVT 1 7 7
TYR HLA-A.02 CLLWSFQTSA 1 7 7
GAD2 HLA-B.08 EAKQKGFVPF 1 5 8
GAD2 HLA-B.08 TLKKMREI 1 5 8
GCN1L1 HLA-A.02 SLLRSLENV 1 0 13
PTPRN HLA-A.02 VIVMLTPLV 1 5 8
PTPRN HLA-B.08 HARIKLKV 1 6 7
ZNT8 HLA-A.01 PTEKGANEY 1 6 7
AKAP9 HLA-A.02 RLSDFSEQL 1 0 12
EXOC8 HLA-A.02 IILVAVPHV 1 0 12
GAD2 HLA-B.08 SRKHKWKL 1 5 7
GANAB HLA-A.02 ALYGFVPVL 1 6 6
GFAP HLA-A.01 ALDIEIATY 1 6 6
INS HLA-A.02 LALWGPDPAA 1 4 7
PABPC1 HLA-A.02 MLGEQLFPL 1 0 11
PTPRN HLA-A.02 MVWESGCTV 1 3 8
GAD2 HLA-A.01 QQDKHYDLSY 1 4 6
GAD2 HLA-B.08 FQQDKHYDL 1 6 4
IAPP HLA-B.08 LNHLKATPI 1 4 6
INSDRIP HLA-A.02 MLYQHLLPL 1 4 6
NDC1 HLA-A.02 CLNEYHLFL 1 0 10
NY-ESO-1 HLA-B.07 APRGPHGGAASGL 2 5 5
PMEL HLA-A.02 ITDQVPFSV 2 2 8
G6PC2 HLA-B.08 NLIFKWKSI 1 5 4
GAD2 HLA-B.08 HPRYFNQL 1 2 7
GAD2 HLA-B.08 MAALPRLIAF 1 5 4
GAD65 HLA-A.02 LLQEYNWEL 1 6 3
IGRP HLA-A.02 YLLLRVLNI 1 2 7
NRAS HLA-A.01 ILDTAGKEEY 1 4 5
CDK4 HLA-A.02 ALDPHSGHFV 1 4 4
G6PC2 HLA-A.01 PTHEEHLFY 1 3 5
GAD65 HLA-A.02 ELAEYLYNI 1 5 3
GNL3L HLA-A.02 NLNCCSVPV 1 4 4
KRAS HLA-C.08 GADGVGKSAL 2 3 5
PPI HLA-A.02 HLCGSHLVEA 1 2 6
PREINS HLA-A.02 HLVEALYLV 1 4 4
RSV-GP HLA-DPA1.01/B1.04 DFHFEVFNFVPCSI 1 4 4
SF3B1 HLA-A.02 ALLPGLPAA 1 4 4
SSX2 HLA-A.02 KASEKIFYV 1 4 4
gp100 HLA-A.02 YLEPGPVTA 2 4 4
p53 HLA-A.02 HMTEVVRHC 2 4 4
G6PC2 HLA-B.08 YVMVTAAL 1 1 6
GAD2 HLA-B.08 RMMEYGTTM 1 4 3
GAD65 HLA-A.02 ILMHCQTTL 1 3 4
IGRP HLA-A.02 FLWSVFMLI 1 2 5
PPM1F HLA-A.02 GVYALIAGA 1 3 4
PTPRN HLA-A.02 SLAAGVKLL 1 3 4
UNC-CDK4-1 HLA-A.02 ALTPVVVTL 1 0 7
ZNT8 HLA-A.02 ILGALLSIL 1 3 4
ZNT8 HLA-A.02 LLIDLTSFL 1 3 4
ZNT8 HLA-A.02 RLLYPDYQI 1 3 4
ASTN1 HLA-A.02 KLYGLDWAEL 1 3 3
GFAP HLA-A.02 NLAQDLATV 1 3 3
GFAP HLA-A.02 QLARQQVHV 1 1 5
IGRP HLA-A.02 FLFAVGFYL 1 2 4
INS HLA-A.02 ALWGPDPAAA 2 2 4
INS HLA-DQA1.03/B1.02 VELGGGPGA 1 3 3
KIF16B HLA-B.07 APARLERRHSA 1 3 3
MMP9 HLA-A.02 NILDAIAEI 1 0 6
PRPF3 HLA-B.27 TRLALIAPK 1 3 3
PTPRN HLA-A.02 ALTAVAEEV 1 3 3
SMARCD3 HLA-A.02 KLFEFLVYGV 2 3 3
TT HLA-DPA1.01/B1.04 FNNFTVSFWLRVPKVSASHLE 1 3 3
WDR46 HLA-A.02 FLIYLDVSV 1 3 3
ZNT8 HLA-A.02 SLNYSGVKEL 1 2 4
p53 HLA-A.68 SSCMGGMNWR 1 3 3
FGFR3 HLA-A.02 SVCAGILSYGV 1 3 2
GAD2 HLA-B.08 YAFLHATDL 1 2 3
GFAP HLA-B.08 EGHLKRNIV 1 2 3
IAPP HLA-B.08 NTYGKRNAV 1 2 3
PGM5 HLA-A.02 AVGSYVYSV 1 2 3
ZNT8 HLA-A.02 LLMEGVPKSL 1 2 3
ZNT8 HLA-A.02 SISVLISAL 1 3 2

Distribution VDJdb confidence scores

Legend: 0 - low confidence, 1 - medium confidence, 2 - high confidence, 3 - very high confidence.

Note that this scoring system is based on generation probabilities acquired via OLGA model. You might apply alternative methods such as TCRNET to infer high-confidence TCR motifs from VDJdb.


Spectratype

Representative data for Homo Sapiens. CDR3 length distribution (spectratype) is colored by cognate epitope. Second plot shows epitope length distribution for MHC class I and II colored by unique CDR3 (alpha or beta) records.

V(D)J usage and MHC alleles

Representative data for Homo Sapiens, Variable gene. Only Variable genes and MHC alleles with at least 10 records are shown.

Circos plot for correspondence between human TRBV genes and MHC class I alleles for links supported by more than 50 records in terms of the ratio of observed to expected records. Band width and color (from red:highest to light yellow:lowest) are scaled proportional to co-occurrence matrix value divided by row (TRBV) and column (HLA) sums.


Detailed summary for HLA

Representative data for Homo Sapiens MHC class I and II

First chain Second chain Records Unique epitopes Studies
HLA-A*02 B2M 31899 653 145
HLA-A*03 B2M 25681 17 13
HLA-A*11 B2M 4726 10 9
HLA-B*08 B2M 4139 67 23
HLA-A*01 B2M 2996 140 19
HLA-B*07 B2M 2614 123 28
HLA-DRA*01 HLA-DRB1*01 1253 15 12
HLA-A*24 B2M 1001 72 16
HLA-E*01 B2M 832 9 4
HLA-B*57 B2M 712 9 8
HLA-B*27 B2M 699 8 13
HLA-DPA*01 HLA-DPB*04 544 1 2
HLA-DRA*01 HLA-DRB1*04 445 13 9
HLA-DRA*01 HLA-DRB5*01 376 8 7
HLA-B*42 B2M 336 7 5
HLA-DRA1*01 HLA-DRB1*15 306 1 1
HLA-B*35 B2M 305 21 21
HLA-DQA1*05 HLA-DQB1*02 305 17 10
HLA-DRA*01 HLA-DRB1*07 295 10 7
HLA-B*44 B2M 291 9 7
HLA-DRA*01 HLA-DRB1*15 242 9 8
HLA-B*15 B2M 233 6 4
HLA-A*68 B2M 212 4 4
HLA-DRA*01 HLA-DRB1*11 189 5 6
HLA-A*08 B2M 117 1 1
HLA-B*40 B2M 112 3 2
HLA-B*18 B2M 80 9 3
HLA-DQA1*03 HLA-DQB1*02 62 9 3
HLA-DQA1*01 HLA-DQB1*06 49 6 6
HLA-B*81 B2M 48 1 2
HLA-C*07 B2M 43 3 2
HLA-DRA*01 HLA-DRB1*03 42 5 3
HLA-DQA1*02 HLA-DQB1*02 32 6 2
HLA-DPA1*01 HLA-DPB1*04 30 8 1
HLA-DRA1*01 HLA-DRB1*11 30 1 1
HLA-B*51 B2M 29 1 1
HLA-DQA*01 HLA-DRB1*07 29 1 1
HLA-DQA1*03 HLA-DQB1*03 26 8 7
HLA-DQA1*01 HLA-DQB1*02 20 1 1
HLA-DQA1*02 HLA-DQB1*06 20 1 1
HLA-DPA1*02 HLA-DPB1*13 16 1 1
HLA-DQA1*05 HLA-DQB1*03 14 5 3
HLA-C*08 B2M 11 3 3
HLA-DQA1*01 HLA-DQB1*05 10 5 4
HLA-B*53 B2M 9 1 1
HLA-DRA*01 HLA-DRB3*03 9 3 3
HLA-B*38 B2M 8 4 2
HLA-A*29 B2M 6 3 1
HLA-DPA1*02 HLA-DPB1*01 6 3 1
HLA-DPA1*02 HLA-DPB1*14 6 3 1
HLA-C*01 B2M 4 1 1
HLA-C*03 B2M 4 2 2
HLA-DPA1*01 HLA-DPB1*02 4 2 2
HLA-DRA*01 HLA-DRB3*02 4 1 1
HLA-A*30 B2M 3 1 1
HLA-DRB1*07 HLA-DRA*01 3 1 1
HLA-A*25 B2M 2 1 1
HLA-B*12 B2M 2 1 1
HLA-B*37 B2M 2 1 1
HLA-B*58 B2M 2 1 1
HLA-C*05 B2M 2 1 1
HLA-C*06 B2M 2 1 1
HLA-C*14 B2M 2 1 1
HLA-DPA1*01 HLA-DPB1*03 2 1 1
HLA-DRA*01 HLA-DRB1*09 2 1 1
HLA-DRA*01 HLA-DRB1*13 2 1 1
HLA-A*32 B2M 1 1 1
HLA-B*14 B2M 1 1 1
HLA-B*52 B2M 1 1 1
HLA-C*04 B2M 1 1 1